University of Groningen
Ruffs in rough times
Schmaltz, Lucie
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2018
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Schmaltz, L. (2018). Ruffs in rough times. University of Groningen.
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The research presented in this thesis was carried out at the Conservation Ecology Group, Groningen Institute for Evolutionary Life Sciences (GELIFES), University of Groningen, The Netherlands.
This research was financially supported by the Ubbo Emmius Fund and the Province of Fryslân.
The printing of this thesis was supported by the University of Groningen and the Province of Fryslân.
COLOFON
Layout and figures; Dick Visser
Cover design: Zsuzsanna Szabó, Lovebird design Photographs: Lucie Schmaltz
Printed by: Lovebird design, Groningen ISBN: 9789403406862
ISBN: 9789403406855 (electronic version) © 2018 Lucie Schmaltz (lucie.schmaltz@gmail.com)
Ruffs in rough times
PhD thesis
to obtain the degree of PhD at the
University of Groningen
on the authority of the
Rector Magnificus Prof. E. Sterken
and in accordance with
the decision by the College of Deans.
This thesis will be defended in public on
Friday 15 June 2018 at 16:15
by
Lucie Emilie Schmaltz
born on 14 May 1984
in Clermont-Ferrand, France
Supervisor
Prof. T. Piersma
Co-supervisor
Prof. J.M. Tinbergen
Assessment committee
Prof. C. Both
Prof. J.A. Gill
Prof. B. Kempenaers
To my grand-mother Alice,
Hoants
Moarnsier It Deel
De dize ús azem
De froast de skou
De dôbers mei iel
De earste ljurk en stirns
Rôpen djerregiel simmerlân
Oer Haklânshop en Kâlde Mage
Hoantsen op ’e wal
It kobbelân, knibbels heech
Mei swietrook earrebarrebrea
Reidlystersang oer lytse weagen
It bûtlân like heech as de Meinesleat
Do wiedest noch hjirre
Wy fiskten de dage mei dy
Us wrâld wie noch oars
Hoantsen op ’e wal
Contents
Chapter 1 General introduction 9
Chapter 2 Quantifying the non-breeding provenance of staging ruffs, Philomachus 23
pugnax, using stable isotope analysis of different tissues
Chapter 3 Apparent annual survival of staging ruffs during a period of population 43 decline: insights from sex and site-use related difference
Chapter 4 Use of agricultural fields by Ruffs staging in southwest Friesland 63 in 2003–2013
Chapter 5 Detection of earthworm prey by Ruff Philomachus pugnax 77
Chapter 6 General discussion 91
References 113
Samenvatting / Friese samenvatting (gearfetting) 131
Acknowledgements 145
List of authors 154
