University of Groningen
Wolves, tree logs and tree regeneration
van Ginkel, Annelies
DOI:
10.33612/diss.112115780
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Publication date:
2020
Link to publication in University of Groningen/UMCG research database
Citation for published version (APA):
van Ginkel, A. (2020). Wolves, tree logs and tree regeneration: Combined effects of downed wood and
wolves on the regeneration of palatable and less palatable tree species. Rijksuniversiteit Groningen.
https://doi.org/10.33612/diss.112115780
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Wolves, tree logs and tree regeneration
Combined effects of downed wood and wolves on the regeneration of
The research presented in this thesis was carried out at the Conservation Ecology Group, Groningen Institute for Evolutionary Life Sciences (GELIFES), University of Groningen, The Netherlands and the Mammal Research Institute of the Polish Academy of Sciences, Poland.
This research is financed by the Ubbo Emmius Fund of the University of Groningen, the KNAW Ecology Fund and by the National Science Center of Poland (grant number 2015/17/B/NZ8/02403).
Pictures: Annelies van Ginkel and Jan van Ginkel Haiku’s: Jan van Ginkel
Layout: Peter van der Sijde, proefschriftgroningen.nl Printed by: Ipskamp
Annelies van Ginkel (hal.van.ginkel@gmail.com) ISBN 978-94-034-2308-1 (book)
ISBN 978-94-034-2309-8 (electronic version)
©2020, H.A.L. van Ginkel. All rights are reserved. No part of this book may be reproduced, distributed, stored in a retrieval system, or transmitted in any form or by any means, without prior written permission of the author.
Wolves, tree logs and tree regeneration
Combined effects of downed wood and wolves on the regeneration of
palatable and less palatable tree species
Proefschrift
ter verkrijging van de graad van doctor aan de
Rijksuniversiteit Groningen
op gezag van de
rector magnificus prof. dr. C. Wijmenga
en volgens besluit van het College voor Promoties.
De openbare verdediging zal plaatsvinden op
vrijdag 7 februari 2020 om 16.15 uur
door
Hermine Annette Lisa van Ginkel
geboren op 17 december 1990
te Westerbork
Promotores
Prof. dr. ir. C. Smit
Prof. dr. D.P.J. Kuijper
Beoordelingscommissie
Prof. dr. ir. F. van Langevelde
Prof. dr. H. Olff
Paranimfen
Emma de Lang
Nadia Hijner
CONTENTS
Chapter 1: General introduction
9
Chapter 2: Wolves and tree logs: landscape-scale and fine-scale risk factors
23
interactively influence tree regeneration
Ecosystems (2019) 22: 202–212
Chapter 3: Impediments affect deer foraging decisions and sapling performance
41
Submitted to Forest Ecology and Management
Chapter 4: Indirect impact of wolf on tree performance is tree species specific and
63
modified by site conditions.
Chapter 5: Safe for saplings not safe for seeds: Quercus robur recruitment in
81
relation to coarse woody debris in Białowieża Primeval Forest, Poland
Forest Ecology and Management (2013) 304;73-79
Chapter 6: Behavioral response of naïve and non-naïve deer to wolf urine
95
PLOS ONE (2019) 14(11): e0223248
Chapter 7: Synthesis
113
Addenda: References
123
List of co-authors
139
Nederlandse samenvatting: wolven, dood hout en bosverjonging
143
Dankwoord / Acknowledgements
155
8
