Het klein drijfhorentje Rissoa parva (da Costa, 1778) van het Banjaardstrand

50 AFZETTINGEN WTKG 40 (2), 2019

De Banjaard

Het klein drijfhorentje Rissoa parva (da Costa, 1778) van het

Banjaardstrand

Han Raven 1

Rissoa parva (da Costa, 1778) from the

Banjaard beach

In a sample of small shells originating from a sand sup-pletion, a huge variation was noted in the specimens of Rissoa parva. Ribbed and smooth forms, as well as in-termediates have been found. The author and Van der Linden (1987) found these forms also together within a single tidal pool in Spain and France, respectively. The extraordinary wide range in forms in the sample from the Banjaard beach is interpreted to be the result of the mixing of material from several strata, mainly Eemian, but possibly also Holocene.

Inleiding

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51 AFZETTINGEN WTKG 40 (2), 2019

JHGRPLQHHUGGRRUKHWZDGVODNMHPeringia ulvae (Pennant, PDDUGHKRRSZDVGDWHURRNDQGHUHNOHLQHVFKHO-SHQLQ]RXGHQ]LWWHQ'DWEOHHN]RWH]LMQRQGHUDQGHUH veel scheefhorens Lacuna vincta 0RQWDJX  YLHU MXYHQLHOH JHQDYHOGH WROKRUHQV Steromphala umbilicalis GD &RVWD  YLMI PHONZLWWH WUDOLHGULMIKRUHQWMHV

Al-vania lactea0LFKDXGHOIYOLH]LJHGULMIKRUHQWMHV Rissoa membranacea -$GDPV  ppQ ZLWWH

ZHQ-teltrap Epitonium clathratulum .DQPDFKHU  GULH VWRPSH buis horentjes Caecum glabrum0RQWDJX HQNHOHPHONZLWWHDUNVFKHOSHQStriarca lactea (Linnaeus, HQppQNOHSMHYDQKHWVFKHHIEXOWVFKHOSMHAltenaeum

dawsoni-HIIUH\VZHUGHQJHYRQGHQQDDVWYHOHDO-JHPHQHVRRUWHQ%RYHQGLHQYHHOH[HPSODUHQYDQKHWNOHLQ drijfhorentje Rissoa parvaGD&RVWD5LDDQ5LMNHQ YRQGRRNHQNHOHH[HPSODUHQYDQObtusella intersecta6 :RRG9ROJHQVKHPLVGLW¿MQHJUXLVPHHVWDOHHQKDOI MDDUQDGHVXSSOHWLHYHUGZHQHQHQEOLMYHQDOOHHQGHJURWHUH VRRUWHQRYHU5LMNHQ

Rissoa parva (da Costa, 1778)

*HHQYDQGHVFKHOSHQYDQGH]HVRRUWZDVYHUV$OOHZDUHQ RQGRRUVFKLMQHQGZLWWRWEOHHNJHHOYDDNPHWUHVWHQYDQKHW NOHXUSDWURRQ1DWXXUOLMN]RXKHWYLQGHQYDQRissoa par-vaRS]LFKQLHWYROGRHQGHDDQOHLGLQJ]LMQYRRUHHQDUWLNHO :DWRSYLHOZDVGHJURWHYHUVFKHLGHQKHLGDDQYRUPHQYDQ GHVFKHOSHQYDQGLWVODNMH ‡ *HULEG¿J$%UHGHVWHYLJHHQYDDNZDWJURWHUHVFKHOS PHWRSGHODDWVWHZLQGLQJRQJHYHHUEUHGHUHFKWHULEEHQ PHWGDDUWXVVHQ¿MQHVSLUDDOVFXOSWXXU0RQGUDQGRPJHVOD-JHQGDDUDFKWHUGHHOVHURSHQHUDFKWHUHHQNRPPDYRUPLJ EUXLQYOHNMH ‡ *ODG¿J'(6ODQNHGXQQHHQYDDNNOHLQHUHVFKHOS ]RQGHU ULEEHQ NOHXUSDWURRQ JHYRUPG GRRU RQJHYHHU  ODQJJHUHNWHRQUHJHOPDWLJHEUXLQHYOHNNHQGLHEHSHUNWNXQ-QHQ]LMQWRWGHDFKWHU]LMGHYDQGHZLQGLQJVRPVRRNRS YRRU]LMGHRIYRRUVWHGHHOHJDDOEUXLQJHNOHXUG0RQGUDQG RPJHVODJHQGDDUDFKWHUGHHOVHURSHQHUDFKWHUHHQNRP-PDYRUPLJEUXLQYOHNMH'H]HYRUPVWRQGODQJEHNHQGDOV DSDUWHVRRUWPDDUZRUGWQXEHVFKRXZGDOVRissoa parva IRUPDinterrupta-$GDPV ‡ 7XVVHQYRUPHQ¿J3B, C, F, G, H'H]H]LMQHULQYHOHYDUL-DWLHV'HULEEHQNXQQHQRSGHODDWVWHZLQGLQJRQWEUHNHQ¿J 3G]HHU]ZDN]LMQRIJHKHHODIZH]LJ¿JB, CGHVFKHOS NDQODQJJHUHNW]LMQDOVLQGHJODGGHYRUPPDDUPHWGHULE- EHQDOVLQGHJHULEGHYRUP¿JHGHULEEHQNXQQHQWDOULM-NHU]LMQLQGLWH[HPSODDURSGHODDWVWHZLQGLQJZDDUELM GHGXQQHJHNURPGHULEEHQLQSDUHQYRRUNRPHQ¿JF 9ROJHQV5LDDQ5LMNHQSHUVFRPPNZDPGHVRRUWYURH- JHUYDNHUYRRULQ=HHODQGPDDUQLHW]RJURIJHULEGDOVLQ¿-JXXU$ Discussie +HWLVDOJHPHHQEHNHQGGDWHUYHUVFKLOOHQGHYRUPHQ]LMQ ELMYRRUEHHOGSUDFKWLJJHwOOXVWUHHUGGRRU6PLWK9HU- VFKLOOHQGHYRUPHQNXQQHQQDDVWHONDDULQHHQHQNHOHJHWLM-GHQSRHOYRRUNRPHQHLJHQZDDUQHPLQJLQ*LMyQ$VWXULDV 6SDQMH¿J$WPF9DQGHU/LQGHQEHYHVWLJWGLW PDDUPHUNWRRNRSGDWLQGH]HOIGHSRHOLQ%UHWDJQHLQKHW YRRUMDDUGHPHHUGHUKHLGYDQGHGLHUHQWRWGHJODGGH YRUPEHKRUHQHQLQKHWQDMDDUWRWGHJHULEGHYRUP :LJKDPPHOGGHDOGDWLQSRSXODWLHVXLWKHW9HUHQLJG .RQLQNULMNLQODWHZLQWHUYURHJHOHQWHGHJODGGHYRUPGR- PLQHHUWHQLQ]RPHUHQQDMDDUGHJHULEGHYRUP,QGHPRQ- VWHUVGLHLQ*LMyQWLMGHQVYHUVFKLOOHQGH]RPHUVZHUGHQJH-QRPHQRYHUKHHUVWHVWHHGVGHJHULEGHYRUPVRPVWRW 'HYDULDWLHELQQHQGHH[HPSODUHQYDQKHW%DQMDDUGVWUDQG LVQRJJURWHUGDQHOGHUV'DWNDQYHUNODDUGZRUGHQGRRUGH KHUNRPVWELMKHWZLQQHQYDQ]DQGYRRUHHQVXSSOHWLHZRUGW YDDNPDWHULDDOXLWYHUVFKLOOHQGHDI]HWWLQJHQJHPHQJG'H NOHXUYDQGHVFKHOSHQJHHIWDDQGDWGLWZDDUVFKLMQOLMNYRRUDO PDWHULDDOXLWKHW(HPLHQEHWUHIWPRJHOLMNXLWYHUVFKLOOHQ-GHODJHQ¿J$&WP HJHPHQJGPHWYHUVHUXLW]LHQG YHUPRHGHOLMN+RORFHHQPDWHULDDO¿JB 2YHULJHQVVFKUHHI9HUGXLQRYHUHHQDQGHUHYDULD-WLHLQVRPPLJHPRQVWHUVYDQKHP]HOIHQERYHQJHQRHPG PRQVWHUYDQ9DQGHU/LQGHQZDUHQGHJHULEGHH[HPSOD-Fig. 2. Vloedlijn met horentjesgruis.

52 AFZETTINGEN WTKG 40 (2), 2019 UHQWHVSOLWVHQLQWZHHJURHSHQJURHS$KDGHHQODQJHUH VFKHOSHHQDQGHUULESDWURRQHQHHQJODGGHUHVFKHOSGDQ GLHLQJURHS%9HUGXLQGDFKWPHWWZHHVRRUWHQWHPDNHQ WHKHEEHQPDDUGLWLVQRRLWYHUGHURSJHYROJG=LMQFROOHF-WLHHQGLHYDQ9DQGHU/LQGHQ]LMQRSJHQRPHQLQ1DWXUDOLV %LRGLYHUVLW\&HQWHUGXVGHEHWUHIIHQGHPRQVWHUVNXQQHQ GDDUZRUGHQEHNHNHQ Dankwoord 0HWKXQHQWKRXVLDVWHYHUKDOHQ)DFHERRNPHOGLQJHQVWLPX- OHHUGHQ5LDDQ5LMNHQHQ+HVWHU/RHIIPLMGLWVWUDQGWHEH-]RHNHQ5LDDQ5LMNHQHQ*HUDUGYDQGHU9HOGHOD]HQKHW YHUKDDORRNGRRUHQPDDNWHQVXJJHVWLHV)UDQN:HVVHOLQJK 1DWXUDOLVZRUGWEHGDQNWYRRUKHWEHVFKLNEDDUVWHOOHQYDQ GHPLFURVFRRSZDDUPHHGHIRWR¶VGRRUPLM]LMQJHPDDNW A B C D E F 1 mm A B C D E F G H 1 mm

Fig.3. Klein drijfhorentje Rissoa parva (da Costa, 1778). Foto’s Han Raven.

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