117
isolate S. yirgalemense. The results obtained also confirmed that most biological control agents cannot be used as a stand-alone control measure, but that they should be applied in combination with other control measures, in an integrated pest management system (IPM). Even 60% control of the overwintering codling moth population, would be sufficient to rid the next generation of those insect larvae escaping chemical control with possible resistance development. However, even though the results from this study clearly show that, under optimum field conditions, and with a highly virulent nematode isolate, good codling moth control can be obtained, combining EPNs with another biocontrol agent, such as entomopathogenic fungi, should be considered in future research.