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The molecular analyses support the distinction of the different species as defined by the morphology and group the morphologically similar C

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(that is, a group with rounded sclerenchyma bundles and a group with elongated sclerenchyma bundles), but further investigations are still needed.

The molecular analyses support the distinction of the different species as defined by the morphology and group the morphologically similar C. fruticosus and C.

squarrosus in close relation. The Boerhavia species are divided into two subclades.

Boerhavia cordobensis, B. diffusa var. diffusa and B. erecta are species alien to southern Africa and form a distinctive subclade. Boerhavia deserticola, B.

hereroensis and B. repens subsp. repens are indigenous to southern Africa and group together in a second subclade. Boerhavia coccinea var. coccinea (whose origin is not known) groups with the indigenous Boerhavia species. The groupings of the Commicarpus species are varied and not ascribable to specific non-molecular trends.

The extra-African C. plumbagineus and C. helenae var. helenae, and the African limited C. pentandrus and C. decipiens group together, but this is not supported by the morphology.

This investigation of the phylogeny was indeed preliminary, as more samples and genes still need to be incorporated and the results interpreted in combination with the morphological, anatomical, palynological and biogeographical data.

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